Seit 12/2024

• Wissenschaftlicher Mitarbeiter (Post-Doc), Institut für Geologische Wissenschaften, Freie Universität Berlin, Deutschland

Seit 12/2024

• Research Associate, Leibniz Institut für die Analyse des Biodiversitätswandels, Hamburg, Deutschland

Seit 04/2017

• Research Associate, Department of Vertebrate Paleontology, American Museum of Natural History, New York City, USA

05/2020 – 11/2024

• Wissenschaftlicher Mitarbeiter (Post-Doc), Fachbereich Biologie, Universität Hamburg, Deutschland

07/2017 – 04/2020

• Theodore Roosevelt Memorial Fund and Division of Paleontology Postdoctoral Fellow, Department of Vertebrate Paleontology, American Museum of Natural History, New York City, USA

05/2015 – 04/2017

• Train2Move Marie Curie Postdoctoral Fellow, Dipartimento di Scienze della Terra, Universita di Torino, Italia

04/2014 – 03/2015

• Volkswagen Stiftung Postdoctoral Fellow, Faculdade de Ciencias e Tecnologia, Universidade NOVA de Lisboa, Caparica, Portugal

01/2010 – 02/2014

• PhD candidate, Faculdade de Ciencias e Tecnologia, Universidade NOVA de Lisboa, Caparica, Portugal

09/2002 – 10/2008

• Bachelor and Master of Science, Fachbereich Biologie, Universität Zürich, Schweiz

Integrative Paläontologie (MSc-PA008)

Lehrbücher:

Nature Through Time: https://link.springer.com/book/10.1007/978-3-030-35058-1

Vertebrate Life: https://global.oup.com/ukhe/product/vertebrate-life-9780197558621?q=vertebrate%20life&cc=de&lang=en

Emanuel is an evolutionary paleobiologist specializing in sauropod dinosaurs and lacertid lizards. His research interests include understanding how sauropods evolved, how they reached their enormous sizes, and how their communities were structured. He also uses lacertid lizards as a model system to investigate intraspecific variation and how this variation can improve phylogenetic inference and species delimitation based on morphology.

Current Research Projects:

• Systematics and phylogeny of diplodocoid sauropods
• Ontogenetic niche differentiation in sauropods from the Morrison Formation, USA
• Soft tissue preservation in the Howe Quarry, a sauropod Lagerstätte in the Morrison Formation of Wyoming
• Phenotypic phylogenetic analysis and species delimitation
• Intraspecific variation in extant and extinct lacertid lizards and how changes in disparity correlate with climate

Publication Highlights

• Tschopp E, O Mateus, RBJ Benson. 2015a. A specimen-level phylogenetic analysis and taxonomic revision of Diplodocidae (Dinosauria, Sauropoda). PeerJ 3: e857.
• Tschopp E, JG Napoli, LCM Wencker, M Delfino, P Upchurch. 2022. How to render species comparable taxonomic units through deep time: a case study on intraspecific osteological variability in extant and extinct lacertid lizards. Systematic Biology, 71:875-900.
• Tschopp E. 2020. Paleontology: the smallest of the giants. Current Biology 30: R1496-R1499.

Papers and Book Chapters, peer reviewed

• Saleiro A*, Tschopp E. 2025. New sauropod dinosaur teeth from the Upper Jurassic of Portugal and their implications for sauropod dental evolution. Papers in Palaeontology 11: e70001.
• Van der Linden T, Wallaard J, Tschopp E, Holwerda F, Schulp A. 2024. A new diplodocid species from the Upper Jurassic Morrison Formation of northern Wyoming. Palaeontologia Electronica 27: a50.
• Lei R*, Tschopp E, Hendrickx C, Wedel MJ, Norell M, Hone DWE. 2023. Bite and tooth marks on sauropod dinosaurs from the Morrison Formation. PeerJ 11:e16327.
• Conti S*, Tschopp E, Mateus O, Zanoni A, Masarati P, Sala G. 2022. Multibody analysis and soft tissue strength refute supersonic dinosaur tail. Scientific Reports 12: 19245.
• Demuth OE*, Benito J, Tschopp E, Lautenschlager S, Mallison H, Heeb N, Field DJ. 2022. Topology-based three-dimensional reconstruction of delicate skeletal fossil remains and the quantification of their taphonomic deformation. Frontiers in Ecology and Evolution 10: 828006.
• Seghetti SM*, Georgalis GL, Tschopp E, Delfino M. 2022. A historical overview of the reptile fauna from the Eocene Bolca Fossil-Lagerstätte (Italy). Bollettino della Società Paleontologica Italiana 61:119-143.
• Tschopp E, Whitlock JA, Woodruff DC, Foster JR, Lei R*, Giovanardi S*. 2022. The Morrison Formation Sauropod Consensus: A freely accessible online spreadsheet of collected sauropod specimens, their housing institutions, contents, references, localities, and other potentially useful information. Peer Community Journal, e17.
• Delfino M, Luján AH, Abella J, Alba DM, Böhme M, Pérez-Ramos A, Tschopp E, Morales J, Montoya P. 2021. Late Miocene remains from Venta del Moro (Iberian Peninsula) provide further insights on the dispersal of crocodiles across the late Miocene Tethys. Journal of Paleontology 95: 184-192.
• Mannion PD, Tschopp E, Whitlock JA. 2021. Anatomy and systematics of the diplodocoid Amphicoelias altus supports high sauropod dinosaur diversity in the Upper Jurassic Morrison Formation of the USA. Royal Society Open Science 8: 210377.
• Seghetti SM*, A Villa*, E Tschopp, F Bernardini, L Laddaga, M Fanelli*, R Levi, M Delfino. 2021. Skull osteology of Vipera walser (Squamata, Viperidae): description, variability, ontogeny, and diagnostic characters in the context of the Italian vipers. Journal of Morphology 282: 5-47.
• Sorbelli L*, A Villa, S Gentili, M Cherin, G Carnevale, E Tschopp, M Delfino.2021. The Early Pleistocene ectothermic vertebrates of Pietrafitta (Italy) and the last European occurrence of Latonia Meyer, 1843. Comptes Rendus Palevol 20: 555-583.
• Wencker LCM*, E Tschopp, A Villa, ML Augé, M Delfino. 2021. Phylogenetic value of jaw elements of lacertid lizards (Squamata: Lacertoidea): a case study with Oligocene material from France. Cladistics 37: 765-802.
• Yu C, Q Jiangzuo, E Tschopp, H Wang, MA Norell. 2021. Information in morphological characters. Ecology and Evolution 11: 11689-11699.
• Fabbri M, E Tschopp, B McPhee, S Nesbitt, D Pol, MC Langer. 2020. Sauropodomorpha F. R. von Huene 1932. In de Queiroz K, P Cantino, JA Gauthier (eds), Phylonyms, a companion to the PhyloCode: 1225-1234. CRC Press, Boca Raton.
• Hendrickx C, E Tschopp, MD Ezcurra. 2020. Taxonomic identification of isolated theropod teeth: the case of the shed tooth crown associated with Aerosteon (Theropoda: Megaraptora) and the dentition of Abelisauridae. Cretaceous Research 108:104312.
• Rio JP, PD Mannion, E Tschopp, JE Martin, M Delfino. 2020. Reappraisal of the morphology and phylogenetic relationships of the alligatoroid crocodylian Diplocynodon hantoniensis from the late Eocene of the United Kingdom. Zoological Journal of the Linnean Society 188: 579-629.
• Tschopp E, C Mehling, MA Norell. 2020. Reconstructing the specimens and history of the Howe Quarry (Upper Jurassic Morrison Formation; Wyoming, USA). American Museum Novitates 3956: 1-56.
• Tschopp E, DE Barta, W Brinkmann, JR Foster, FM Holwerda, SCR Maidment, SF Poropat, TM Scheyer, AG Sellés, B Vila, M Zahner. 2020. How to live with dinosaurs: Ecosystems across the Mesozoic. In Martinetto E, Tschopp E, Gastaldo RA (eds), Nature through Time: 209-230. Springer, Switzerland.
• Tschopp E, R Araújo, SL Brusatte, C Hendrickx, L Macaluso*, SCR Maidment, M Rabi, D Rashid, C Romano, T Williamson. 2020. Dinosaurs, but not only: Vertebrate evolution throughout the Mesozoic. In Martinetto E, Tschopp E, Gastaldo RA (eds), Nature through Time: 187-208. Springer, Switzerland.
• Tschopp E, P Upchurch. 2019. The challenges and potential utility of phenotypic specimen-level phylogeny based on maximum parsimony. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 109: 301-323.
• Tschopp E, SCR Maidment, MC Lamanna, MA Norell. 2019. Reassessment of a historical collection of sauropod dinosaurs from the northern Morrison Formation of Wyoming, with implications for sauropod biogeography. Bulletin of the American Museum of Natural History 437: 1-79.
• Macaluso L*, E Tschopp. 2018. Evolutionary changes in pubic orientation in dinosaurs are more strongly correlated with the ventilation system than with herbivory. Palaeontology 61:703-719.
• Maltese A, E Tschopp, FM Holwerda, D Burnham. 2018. The real Bigfoot: a pes from Wyoming, USA is the largest sauropod pes ever reported and the northern-most occurrence of brachiosaurids in the Upper Jurassic Morrison Formation. PeerJ 6:e5250.
• Tschopp E, A Villa, M Camaiti*, L Ferro*, C Tuveri, L Rook, M Arca, M Delfino. 2018. The first fossils of Timon (Squamata: Lacertinae) from Sardinia (Italy) and potential causes for its local extinction in the Pleistocene. Zoological Journal of the Linnean Society 184 (3): 825-856.
• Tschopp E, D Brinkman, J Henderson, MA Turner, O Mateus. 2018. Considerations on the replacement of a type species in the case of the sauropod dinosaur Diplodocus Marsh, 1878. Geology of the Intermountain West 5:245-262.
• Tschopp E, FA Tschopp, O Mateus. 2018. Overlap Indices: Tools to quantify the amount of anatomical overlap among groups of incomplete terminal taxa in phylogenetic analyses. Acta Zoologica 99:169–176.
• Tschopp E, O Mateus, MA Norell. 2018. Complex overlapping joints between facial bones allowing limited anterior sliding movements of the snout in diplodocid sauropods. American Museum Novitates 3911:1-16.
• Mateus S, E Tschopp. 2017. Scientific illustration and reconstruction of a skull of the diplodocid sauropod dinosaur Galeamopus. Journal of Paleontological Techniques 17:1–11.
• Tschopp E, O Mateus. 2017. Osteology of Galeamopus pabsti sp. nov. (Sauropoda: Diplodocidae), with implications for neurocentral closure timing, and the cervico-dorsal transition in diplodocids. PeerJ 5:e3179.
• Villa A, E Tschopp, GL Georgalis, M Delfino. 2017. Osteology, fossil record and palaeodiversity of the European lizards. Amphibia-Reptilia 38:79–88.
• Ceríaco LMP, EE Gutiérrez, A Dubois, et al. (in total 493 co-authors). 2016. Photography-based taxonomy is inadequate, unnecessary, and potentially harmful for biological sciences. Zootaxa 4196:435–445.
• Tschopp E. 2016. Nomenclature of vertebral laminae in lizards, with comments on ontogenetic and serial variation in Lacertini (Squamata, Lacertidae). PLoS ONE 11 (2): e0149445.
• Tschopp E, O Mateus. 2016. Case 3700: Diplodocus Marsh, 1878 (Dinosauria, Sauropoda): proposed designation of D. carnegii Hatcher, 1901 as the type species. Bulletin of Zoological Nomenclature 73(1): 17-24.
• Tschopp E, O Wings, T Frauenfelder, BM Rothschild. 2016. Pathological phalanges in a camarasaurid sauropod dinosaur and implications on behaviour. Acta Palaeontologica Polonica 61 (1): 125-134.
• Villa A, Z Roček, E Tschopp, LW van den Hoek Ostende, M Delfino. 2016. Palaeobatrachus eurydices, sp. nov. (Amphibia, Anura), the last western European palaeobatrachid. Journal of Vertebrate Paleontology 36:e1211664.
• Tschopp E, O Wings, T Frauenfelder, W Brinkmann. 2015b. Articulated bone sets set of manus and pedes of Camarasaurus (Sauropoda, Dinosauria). Palaeontologia Electronica 18.2.44A: 1-65.
• Tschopp E, O Mateus. 2013a. Clavicles, interclavicles, gastralia, and sternal ribs in sauropod dinosaurs: new reports from Diplodocidae and their morphological, functional, and evolutionary implications. Journal of Anatomy 222:321–340.
• Tschopp E, O Mateus. 2013b. The skull and neck of a new flagellicaudatan sauropod from the Morrison Formation and its implication on evolution and ontogeny of diplodocid dinosaurs. Journal of Systematic Palaeontology 11:853–888.
• Tschopp E, J Russo*, G Dzemski. 2013. Retrodeformation as a test for the validity of phylogenetic characters: an example from diplodocid sauropod vertebrae. Palaeontologia Electronica 16:1–23.
• Tschopp E, G Dzemski. 2012. 3-dimensional reproduction techniques to preserve and spread paleontological material. Journal of Paleontological Techniques 10: 1-8.
• Christiansen NA, E Tschopp. 2010. Exceptional stegosaur integument impressions from the Upper Jurassic Morrison Formation of Wyoming. Swiss Journal of Geosciences 103: 163-171.

Edited Volumes

• Martinetto E, E Tschopp, RA Gastaldo (eds.) 2020. Nature through Time: Virtual field trips through the Nature of the past. Springer Textbooks, Springer Verlag, Switzerland. 462pp.
• Tschopp E (ed.) 2019. Book Reviews, volume 3. Journal of Paleontological Techniques, book reviews 3. 14pp.
• Tschopp E, F Holwerda, S Costa (eds.) 2018. Book Reviews, volume 2. Journal of Paleontological Techniques, book reviews 2. 7pp.
• Tschopp E, S Costa (eds.) 2017. Book Reviews, volume 1. Journal of Paleontological Techniques, book reviews 1. 6pp.
• Tschopp E, F Holwerda, S Val (eds.) 2016. Symposium Volume, 2^nd International Conservation Symposium Workshop of Natural History Collections. Journal of Paleontological Techniques 15. 161pp.
• Tschopp E, S Val (eds.) 2014. Symposium Volume, 1^st International Conservation Symposium Workshop of Natural History Collections. Journal of Paleontological Techniques 13. 117pp.

Book chapters, not peer-reviewed

• Tschopp E. 2022. Avemetatarsalia and the origin of Dinosauria. In Pough FH, Bemis WE, McGuire B, Janis CM (eds), Vertebrate Life: 397-420. Oxford University Press, New York, USA.
• Tschopp E. 2022. Theropods and the origin of Birds. In Pough FH, Bemis WE, McGuire B, Janis CM (eds), Vertebrate Life: 421-442. Oxford University Press, New York, USA.

Popular Science articles

• Tschopp E. 2017. Brontosaurus und Europasaurus – Sauropoden waren nicht alle gigantisch. Pp. 32-37 in: A. Beckmann, C. Hühne, and R. Kosma (eds.), Jurassic Harz. Dinosaurier von Oker bis Wyoming. Verlag Dr. Friedrich Pfeil, München. 116 pp.
• Tschopp E. 2014. Was clevere Delfine von ihrer Mutter lernen. Tagesanzeiger 93: 32.
• Tschopp E. 2012. Dinosaurier: Woher Forscher wissen, wie sie aussahen. Spick, das schlaue Schülermagazin aus der Schweiz 359: 6-10.
• Tschopp E. 2011. Gefieder der Urvögel in 3-D. Neue Zürcher Zeitung220: 59.
• Tschopp E. 2011. Autopsie an 110 Millionen Jahre altem Dinosaurier. Neue Zürcher Zeitung155: 58.
• Tschopp E. 2011. Mit nur ein, zwei Knochen zu einem neuen Dinosaurier. Neue Zürcher Zeitung 97: 58.

PrePrints

• Cui Q, Yu C, Ma C, Ren Q, Tschopp E, Pol D, Chen Y. 2024. Monsoon Shaped Herbivore Dinosaur Distribution. Available at SSRN: http://dx.doi.org/10.2139/ssrn.4750442
• Saleiro A*, Tschopp E, Daasch A, Kaiser TM, Wiesinger R, Winkler DE. 2024. Acquisition of a dataset for dental microwear analysis (DMTA) of sauropod teeth: pre-analytical and post-processing steps for quality assessment. PaleorXiv. https://doi.org/10.31233/osf.io/8uhsp
• Yu C, Jiangzuo Q, Tschopp E, Wang H, Norell MA. 2021. Information in morphological characters. Authorea. https://doi.org/10.22541/au.162281273.31812918/v1
• Tschopp E,Whitlock LA, Woodruff DC, Foster JR, Lei R*, Giovanardi S*.2019. The Morrison Formation sauropod consensus: A freely accessible online spreadsheet of collected sauropod specimens, their housing institutions, contents, references, localities, and other potentially useful information. PaleorXiv. https://doi.org/10.31233/osf.io/fy43t

Book Reviews

• Tschopp E. 2019. Fossilien im Alpstein, edited by Peter Kürsteiner and Christian Klug (2018). Journal of Paleontological Techniques, book review 3: 12-13.
• Tschopp E. 2017. Species + Speciation in the Fossil Record, edited by Warren D. Allmon and Margareth M. Yacobucci (2016). Journal of Paleontological Techniques, book review 1: 5-6.
• Tschopp E. 2014. Dinosaur Paleobiology, by Stephen Brusatte (2012). The Palaeontological Association Newsletter 85: 85-86.